Enables metallocarboxypeptidase activity and tubulin binding activity. Involved in C-terminal protein deglutamylation; defense response to virus; and protein side chain deglutamylation. Acts upstream of or within several processes, including axonal transport of mitochondrion; positive regulation of ubiquitin-dependent protein catabolic process; and regulation of blastocyst development. Located in cytosol. Is expressed in brain; nose; and trigeminal ganglion. Orthologous to human AGBL4 (AGBL carboxypeptidase 4). [provided by Alliance of Genome Resources, Apr 2022]