Enables sphingosine N-acyltransferase activity. Acts upstream of or within ceramide biosynthetic process and negative regulation of fat cell differentiation. Located in Golgi apparatus and endoplasmic reticulum. Is expressed in nervous system and neural retina. Human ortholog(s) of this gene implicated in cone-rod dystrophy. Orthologous to human TLCD3B (TLC domain containing 3B). [provided by Alliance of Genome Resources, Apr 2022]