Enables promoter-specific chromatin binding activity. Involved in several processes, including aging; positive regulation of smooth muscle cell migration; and response to water-immersion restraint stress. Located in cytosol and nucleus. Biomarker of alcohol-induced mental disorder; gastric ulcer; and paraplegia. Orthologous to human FOXO4 (forkhead box O4). [provided by Alliance of Genome Resources, Apr 2022]