Enables several functions, including ATPase binding activity; GTP binding activity; and ionotropic glutamate receptor binding activity. Involved in Rab protein signal transduction. Located in cytoplasmic vesicle; cytosol; and plasma membrane. Is anchored component of synaptic vesicle membrane. Orthologous to human RAB4A (RAB4A, member RAS oncogene family). [provided by Alliance of Genome Resources, Apr 2022]