Acts upstream of or within spermatid development. Located in condensed nuclear chromosome; cytoplasm; and cytoskeleton. Is expressed in several structures, including alimentary system; brain; genitourinary system; sensory organ; and skeleton. Human ortholog(s) of this gene implicated in primary ciliary dyskinesia 24. Orthologous to human RSPH1 (radial spoke head component 1). [provided by Alliance of Genome Resources, Apr 2022]