Enables several functions, including actin filament binding activity; calmodulin binding activity; and protein kinase C binding activity. Involved in actin crosslink formation and actin filament bundle assembly. Located in several cellular components, including cell cortex; cytoskeleton; and germinal vesicle. Is expressed in several structures, including embryo ectoderm; head skin; nervous system; sensory organ; and tongue mesenchyme. Orthologous to human MARCKS (myristoylated alanine rich protein kinase C substrate). [provided by Alliance of Genome Resources, Apr 2022]