Enables L-proline transmembrane transporter activity. Involved in proline transport. Is active in Schaffer collateral - CA1 synapse and glutamatergic synapse. Is integral component of presynaptic membrane and integral component of synaptic vesicle membrane. Orthologous to human SLC6A7 (solute carrier family 6 member 7). [provided by Alliance of Genome Resources, Apr 2022]