Enables L-histidine transmembrane transporter activity and peptide:proton symporter activity. Involved in several processes, including L-histidine transmembrane export from vacuole; positive regulation of pattern recognition receptor signaling pathway; and regulation of isotype switching to IgG isotypes. Located in early endosome membrane and lysosomal membrane. Is integral component of endosome membrane and integral component of lysosomal membrane. Is expressed in embryo. Orthologous to human SLC15A4 (solute carrier family 15 member 4). [provided by Alliance of Genome Resources, Apr 2022]