Enables ATP hydrolysis activity and TBP-class protein binding activity. Involved in modulation of chemical synaptic transmission and positive regulation of inclusion body assembly. Located in inclusion body; nucleus; and postsynapse. Part of cytosolic proteasome complex and nuclear proteasome complex. Orthologous to human PSMC5 (proteasome 26S subunit, ATPase 5). [provided by Alliance of Genome Resources, Apr 2022]