Enables several functions, including SNAP receptor activity; small GTPase binding activity; and spectrin binding activity. Involved in several processes, including exocytosis; membrane fusion; and protein localization to cell surface. Located in several cellular components, including basolateral plasma membrane; neuron projection membrane; and sarcolemma. Part of SNARE complex. Is active in glutamatergic synapse. Orthologous to human STX4 (syntaxin 4). [provided by Alliance of Genome Resources, Apr 2022]