Enables metallocarboxypeptidase activity and tubulin binding activity. Involved in several processes, including nervous system development; neuromuscular process; and protein deglutamylation. Acts upstream of or within several processes, including adult walking behavior; axonal transport of mitochondrion; and central nervous system neuron differentiation. Located in cytosol; mitochondrion; and nucleus. Is expressed in central nervous system; dorsal root ganglion; and retina. Used to study retinitis pigmentosa. Orthologous to human AGTPBP1 (ATP/GTP binding carboxypeptidase 1). [provided by Alliance of Genome Resources, Apr 2022]