Enables ATP hydrolysis activity and proton-transporting ATPase activity, rotational mechanism. Predicted to be involved in proton transmembrane transport. Predicted to act upstream of or within ion transport. Is integral component of synaptic vesicle membrane. Part of vacuolar proton-transporting V-type ATPase complex. Is expressed in several structures, including nervous system. Orthologous to human ATP6V1G2 (ATPase H+ transporting V1 subunit G2). [provided by Alliance of Genome Resources, Apr 2022]