Enables dynein intermediate chain binding activity; protein C-terminus binding activity; and protein domain specific binding activity. Acts upstream of or within intraciliary retrograde transport; motile cilium assembly; and positive regulation of non-motile cilium assembly. Located in cytoskeleton; cytosol; and membrane. Part of cytoplasmic dynein complex. Is expressed in several structures, including central nervous system; heart; liver; lung; and sensory organ. Orthologous to human DYNLL1 (dynein light chain LC8-type 1). [provided by Alliance of Genome Resources, Apr 2022]