Predicted to enable ATP-dependent activity, acting on DNA. Acts upstream of or within gamete generation; homologous chromosome pairing at meiosis; and ovarian follicle development. Located in recombination nodule and synaptonemal complex. Is expressed in several structures, including brain; gonad; heart; and thymus. Orthologous to human MSH4 (mutS homolog 4). [provided by Alliance of Genome Resources, Apr 2022]