Enables sequence-specific DNA binding activity. Acts upstream of or within mitotic G2 DNA damage checkpoint signaling; negative regulation of cell population proliferation; and positive regulation of transcription, DNA-templated. Predicted to be located in cytosol and nuclear speck. Predicted to be active in nucleus. Is expressed in several structures, including central nervous system; early embryo; genitourinary system; inner cell mass; and sensory organ. Orthologous to human FOXO4 (forkhead box O4). [provided by Alliance of Genome Resources, Apr 2022]