Enables dynein complex binding activity and identical protein binding activity. Involved in cytoskeleton organization; spindle localization; and telomere localization. Acts upstream of or within double-strand break repair via homologous recombination; gamete generation; and homologous chromosome segregation. Located in chromosome; meiotic spindle pole; and nuclear outer membrane. Part of meiotic nuclear membrane microtubule tethering complex. Is expressed in spermatocyte. Orthologous to human KASH5 (KASH domain containing 5). [provided by Alliance of Genome Resources, Apr 2022]