Enables mitogen-activated protein kinase kinase kinase binding activity. Involved in cerebellar cortex development; positive regulation of JNK cascade; and positive regulation of axonogenesis. Located in axon terminus; cytoplasm; and neuronal cell body. Part of protein-containing complex. Orthologous to human DIXDC1 (DIX domain containing 1). [provided by Alliance of Genome Resources, Apr 2022]