Enables MAP-kinase scaffold activity and mitogen-activated protein kinase kinase binding activity. Involved in positive regulation of cold-induced thermogenesis. Acts upstream of or within calcium-mediated signaling and positive regulation of MAPK cascade. Located in cytosol and plasma membrane. Orthologous to human KSR2 (kinase suppressor of ras 2). [provided by Alliance of Genome Resources, Apr 2022]