Enables DNA-binding transcription factor activity and peroxisome proliferator activated receptor binding activity. Involved in cellular response to ionomycin and regulation of synaptic plasticity. Located in cytosol and nucleus. Human ortholog(s) of this gene implicated in dilated cardiomyopathy. Orthologous to human NFATC4 (nuclear factor of activated T cells 4). [provided by Alliance of Genome Resources, Apr 2022]