Enables actin filament binding activity and cell adhesion molecule binding activity. Involved in several processes, including Cdc42 protein signal transduction; actin filament bundle retrograde transport; and netrin-activated signaling pathway. Acts upstream of or within axonogenesis. Located in axonal growth cone; filopodium; and lamellipodium. Orthologous to human SHTN1 (shootin 1). [provided by Alliance of Genome Resources, Apr 2022]