Enables actin binding activity. Acts upstream of or within several processes, including columnar/cuboidal epithelial cell development; neural tube closure; and regulation of cell shape. Located in several cellular components, including adherens junction; apical junction complex; and apical plasma membrane. Is expressed in several structures, including central nervous system; gut; paraxial mesenchyme; sensory organ; and skeletal musculature. Orthologous to human SHROOM3 (shroom family member 3). [provided by Alliance of Genome Resources, Apr 2022]