Enables G-protein beta/gamma-subunit complex binding activity; heterotrimeric G-protein binding activity; and microtubule binding activity. Acts upstream of or within spermatogenesis. Located in several cellular components, including 9+2 motile cilium; microtubule organizing center; and photoreceptor connecting cilium. Is expressed in 4th ventricle lateral recess; choroid plexus; and lateral ventricle choroid plexus. Orthologous to human CETN2 (centrin 2). [provided by Alliance of Genome Resources, Apr 2022]