Predicted to enable proteasome-activating activity. Acts upstream of or within blastocyst development. Part of proteasome accessory complex. Is expressed in several structures, including alimentary system; cranial ganglion; early conceptus; gonad; and liver lobe. Used to study amyotrophic lateral sclerosis. Orthologous to human PSMC4 (proteasome 26S subunit, ATPase 4). [provided by Alliance of Genome Resources, Apr 2022]