Enables ubiquitin protein ligase activity. Involved in several processes, including histone H2A ubiquitination; male gamete generation; and negative regulation of transposition. Acts upstream of or within male meiosis I and ubiquitin-dependent protein catabolic process. Part of ubiquitin ligase complex. Colocalizes with chromatin. Is expressed in several structures, including alimentary system; endocrine gland; eye; genitourinary system; and trachea. Orthologous to human UBR2 (ubiquitin protein ligase E3 component n-recognin 2). [provided by Alliance of Genome Resources, Apr 2022]