Enables cysteine-type endopeptidase inhibitor activity involved in apoptotic process. Involved in several processes, including cellular response to hepatocyte growth factor stimulus; intestinal D-glucose absorption; and regulation of plasma membrane bounded cell projection organization. Located in several cellular components, including brush border; filopodium tip; and lamellipodium. Is expressed in several structures, including brain; egg cylinder; gut; metanephros; and reproductive system. Orthologous to human VIL1 (villin 1). [provided by Alliance of Genome Resources, Apr 2022]