Enables tropomyosin binding activity. Involved in lens fiber cell development. Acts upstream of or within adult locomotory behavior; muscle contraction; and myofibril assembly. Located in cortical cytoskeleton; membrane; and sarcomere. Is expressed in several structures, including ganglia; heart; hemolymphoid system; musculature; and sensory organ. Used to study dilated cardiomyopathy. Orthologous to human TMOD1 (tropomodulin 1). [provided by Alliance of Genome Resources, Apr 2022]