Predicted to enable calcium ion binding activity; protein-glutamine gamma-glutamyltransferase activity; and structural molecule activity. Predicted to be involved in cell envelope organization and peptide cross-linking. Predicted to act upstream of or within keratinization. Located in collagen-containing extracellular matrix. Is expressed in several structures, including alimentary system; genitourinary system; integumental system; musculoskeletal system; and nervous system. Orthologous to human TGM3 (transglutaminase 3). [provided by Alliance of Genome Resources, Apr 2022]