Enables signaling receptor binding activity. Acts upstream of or within negative regulation of transcription, DNA-templated. Part of proteasome regulatory particle. Is expressed in several structures, including gonad; hemolymphoid system gland; limb; liver; and lung. Orthologous to human PSMC5 (proteasome 26S subunit, ATPase 5). [provided by Alliance of Genome Resources, Apr 2022]