Enables DNA-binding transcription activator activity, RNA polymerase II-specific; chromatin binding activity; and cis-regulatory region sequence-specific DNA binding activity. Acts upstream of or within several processes, including animal organ development; branching involved in blood vessel morphogenesis; and cellular response to metal ion. Located in cytosol and nucleus. Is expressed in several structures, including alimentary system; central nervous system; genitourinary system; hemolymphoid system gland; and lung. Orthologous to human NFATC3 (nuclear factor of activated T cells 3). [provided by Alliance of Genome Resources, Apr 2022]