Enables GTPase activator activity and microfilament motor activity. Acts upstream of or within several processes, including Rho protein signal transduction; lamellipodium morphogenesis; and leukocyte chemotaxis. Located in filopodium tip; lamellipodium; and ruffle. Is expressed in brain; cerebellum; heart; inner ear; and skeletal muscle. Human ortholog(s) of this gene implicated in celiac disease. Orthologous to human MYO9B (myosin IXB). [provided by Alliance of Genome Resources, Apr 2022]