Enables L-lactate dehydrogenase activity. Acts upstream of or within ATP biosynthetic process; flagellated sperm motility; and lactate metabolic process. Located in cytoplasm and motile cilium. Is expressed in several structures, including alimentary system; brain; genitourinary system; respiratory system; and sensory organ. Orthologous to human LDHC (lactate dehydrogenase C). [provided by Alliance of Genome Resources, Apr 2022]