Enables glycolipid binding activity. Involved in positive regulation of ATPase activity; positive regulation of ATPase-coupled calcium transmembrane transporter activity; and spermatogenesis. Acts upstream of or within several processes, including male gamete generation; positive regulation of G2/M transition of mitotic cell cycle; and synaptonemal complex disassembly. Located in cell surface; meiotic spindle; and nucleus. Part of CatSper complex. Is expressed in alimentary system; ganglia; medulla oblongata basal plate; testis; and vibrissa. Orthologous to human HSPA2 (heat shock protein family A (Hsp70) member 2). [provided by Alliance of Genome Resources, Apr 2022]