Enables NAD+ ADP-ribosyltransferase activity and NAD+ nucleosidase activity. Acts upstream of or within protein ADP-ribosylation. Located in extracellular region and membrane. Is expressed in several structures, including heart; limb; male reproductive system; nose; and skeletal musculature. Orthologous to human ART5 (ADP-ribosyltransferase 5). [provided by Alliance of Genome Resources, Apr 2022]