Enables actin binding activity. Involved in several processes, including modulation of chemical synaptic transmission; regulation of actin filament bundle assembly; and spine apparatus assembly. Located in several cellular components, including Schaffer collateral - CA1 synapse; spine apparatus; and stress fiber. Is active in glutamatergic synapse. Is expressed in several structures, including genitourinary system; respiratory system; sensory organ; skeleton; and tooth. Orthologous to human SYNPO (synaptopodin). [provided by Alliance of Genome Resources, Apr 2022]